- Fukuda Asuka (Formation - April 18, 1999)
- Ichii Sayaka (April 19, 1999 - August 21, 1999)
- Goto Maki (August 22, 1999 - April 15, 2000)
- Kago Ai (April 16, 2000 - August 25, 2001)
- Niigaki Risa (August 26, 2001 - January 18, 2003)
- Tanaka Reina (January 19, 2003 - April 30, 2005)
- Kusumi Koharu (May 1, 2005 - December 9, 2006)
- Mitsui Aika (December 10, 2006 - January 1, 2011)
- Suzuki Kanon (January 2, 2011 - September 28, 2011)
- Kudo Haruka (September 29, 2011 - September 29, 2014)
- Haga Akane (September 30, 2014 - June 21, 2019)
- Yamazaki Mei (June 22, 2019 - June 28, 2022)
- Sakurai Rio (June 29, 2022 - May 22, 2023)
- Yumigeta Ako (May 23, 2023 - present)
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